The environments are stripped of texture and are replaced by simple colour palettes that change as the player reaches new zones. The audio/visual atmosphere during Zone mode differs greatly from regular play. As the player's craft automatically passes through higher 'zones', or categories of speed, the player must continue to navigate the course until their energy runs out and the craft explodes. Finally, the unique Zone mode, first introduced in Wipeout Fusion, automates the player's acceleration control, progressively moving the craft at higher speeds.

In the Speed Lap and Time Trial modes, the player races alone in an attempt to beat the clock. Single Race is a straight-forward start-to-finish race against seven other competitors, while Tournament is a series of consecutive races won by having the highest aggregate score. There are five main race types in Wipeout HD: Single Race, Tournament, Speed Lap, Time Trial, and Zone. The player pilots an anti-gravity craft, selected from one of several teams and, depending on the game mode, competes using speed and weaponry in an attempt to beat the competition.

Gameplay is largely similar to that of previous Wipeout titles. Official logo of the FX350 Anti-Gravity Racing League

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